An engine-driven centrifugal pump accommodated in the engine circulates coolant through the system. The pump takes in the coolant from the bottom section of the radiator and directs it through the ducts into the cylinders and cylinder heads to cool down engine internals.
When coolant expands from heat, the expansion reservoir takes up excess coolant.
"LOW" and "FULL" level marks facilitate coolant level inspection and top-up; see 2.10.1.